the males also

have breasts, like the females, exceedingly small. The she-bear has
four breasts. Some animals have two breasts, but situated near the
thighs, and teats, likewise two in number, as the sheep; others have
four teats, as the cow. Some have breasts neither in the chest nor
at the thighs, but in the belly, as the dog and pig; and they have a
considerable number of breasts or dugs, but not all of equal size.
Thus the shepard has four dugs in the belly, the lioness two, and
others more. The she-camel, also, has two dugs and four teats, like
the cow. Of solid-hooved animals the males have no dugs, excepting
in the case of males that take after the mother, which phenomenon is
observable in horses.